This page is dedicated to cycad-related research. At the top of the page are listed the research grants that have been funded by TCS since our active grant program began in 2007. Below that are some recent examples of cycad research being conducted around the world. Click here for more information about the TCS Grants program.

Research Grants Funded by TCS

2010

• Cycas micronesica on the Island of Yap

2009

• Travel Scholarship to Study Cycads and Associated Flora in Queensland, Australia

• Human Geography and Ethnobotany of Cycads in Xi’iuy, Teenek, and Nahuatl communities of northeastern Mexico

2008

• Cycas debaoensis International Conservation Project

• Recruitment dynamics of Zamia obliqua populations and implications for management and conservation

• Cycas Conservation Centers in the Nilgiri Biosphere Reserve, Western Ghats, India

2007

• Population status and life history of Zamia in the Chocó-Colombia hotspot

• The chamal (Dioon edule Lindl.) in the state of San Luis Potosí

• Conservation of the endemic over-exploited cycad, Cycas circinalis L., in the Nilgiri Biosphere Reserve, South India

Recent Research Articles

2010

• Marler, T.E. 2010. Cycad mutualist offers more than pollen transport. American Journal of Botany 95:1-5.

• Lopez-Gallego, C. & P. O'Neil. 2010. Life-history variation following habitat degradation associated with differing fine-scale spatial genetic structure in a rainforest cycad. Population Ecology 52:191–201.

2009

• Calonje, M. 2009. A new cliff-dwelling species of Zamia (Zamiaceae) from Belize. Journal of the Botanical Research Institute of Texas 3:23-29.

• Calonje, M. & J. Meerman. 2009. What is Zamia prasina (Zamiaceae: Cycadales)? Journal of the Botanical Research Institute of Texas 3:43-49.

• Calonje, M., J. Meerman, P. Griffith & G. Hoese. 2009. A new species of Zamia (Zamiaceae) from the Maya mountains of Belize. Journal of the Botanical Research Institute of Texas 3:31-41.

• Osborne, R., D.W. Stevenson & A.P. Vovides. 2009. What is Ceratozamia fuscoviridis? Delpinoa 48:5-10. (4 Mb)

• Nicolalde-Morejón, F., F. Vergara-Silva, J. González-Astorga, A.P. Vovides & A.E. de los Monteros. 2009. Reciprocal illumination of morphological characters upon a molecular hypothesis supports the proposal of a new species of cycad from Mexico. Systematics and Biodiversity 7:73-79. (12 Mb)

2008

• Haynes, J.L., L.M. Whitelock, B. Schutzman & R.S. Adams. 2008. A new endemic Ceratozamia from Honduras (Cycadales: Zamiaceae). Cycad Newsletter 31(2/3):16-21.

• Schutzman, B., R.S. Adams, J.L. Haynes & L.M. Whitelock. 2008. A new endemic Zamia from Honduras (Cycadales: Zamiaceae). Cycad Newsletter 31(2/3):22-26.

• Taylor, A.S., J.L. Haynes, G. Holzman. 2008. Taxonomical, nomenclatural, and biogeographical revelations in the Zamia skinneri species complex (Cycadales: Zamiaceae) of Central America. Botanical Journal of the Linnean Society 158:399-429.

• Zgurski, J.M., H.S. Rai, Q.M. Fai, D.J. Bogler, J. Francisco-Ortega & S.W. Graham. 2008. How well do we understand the overall backbone of cycad phylogeny? New insights from a large, multigene plastid data set. Molecular Phylogenetics & Evolution, in press.

2007

• Haynes, J. L. & M. A. Bonta. 2007. An emended description of Dioon mejiae Standl. & L.O. Williams (Zamiaceae). Proceedings of the 7th International Conference on Cycad Biology, Xalapa, Mexico, January 2005. Memoirs of the New York Botanical Garden 97: 418-443. (2.1 Mb)

• Marler, T. E., V. Lee, and C. A. Shaw. 2007. Storage does not influence steryl glucoside concentration in Cycas micronesica seeds. HortScience 42(3):626-628.

• Pócs, T. & Q. Luke. 2007. A new species of Encephalartos (Zamiaceae) from Tanzania. Journal of East African Natural History 96:193-201.

• Saneesh, C.S. & A. Varghese. 2007. Mutualistic relationships involving the endemic Cycas circinalis L.: Field notes from the Appankappu Forests, Nilambur, Kerala, India. Cycad Newsletter 30(4):28-29.

• Taylor, A.S., J.L. Haynes, G. Holzman & J. Mendieta. 2007. Variability and structure of natural populations and implications for Zamia species in western Panama. Proceedings of the 7th International Conference on Cycad Biology, Xalapa, Mexico, January 2005. Memoirs of the New York Botanical Garden 97:557-577.

• Terry, I., G. H. Walter, C. Moore, R. Roemer & C. Hull. 2007. Odor-mediated push-pull pollination in cycads. Science 318:70.

2006

• Bonta, M. A., O. Flores Pinot, D. Graham, J. Haynes & G. Sandoval. 2006. Ethnobotany and conservation of tiusinte (Dioon mejiae Standl. & L.O. Williams, Zamiaceae) in Northeastern Honduras. Journal of Ethnobiology 26(2):228-257. (2.6 Mb)

• Fisher, J. B., and T. E. Marler. 2006. Eccentric growth but no compression wood in a horizontal stem of Cycas micronesica (Cycadales). IAWA Journal 27(4):377-382.

• Marler, T. E., V. Lee, J. Chung, and C. A. Shaw. 2006. Steryl glucoside concentration declines with Cycas micronesica seed age. Functional Plant Biology 33:857-862.

• Marler, T. E., and R. Muniappan. 2006. Pests of Cycas micronesica leaf, stem, and male reproductive tissues with notes on current threat status. Micronesica 39(1):1–9.

• Niklas, K. J., E. D. Cobb, and T. Marler. 2006. A comparison between the record height-to-stem diameter allometries of pachycaulis and leptocaulis species. Annals of Botany 97:79-83.

• Pérez-Farrera, M.A. & A.P. Vovides. 2006. The ceremonial use of the threatened “espadaña” cycad (Dioon merolae, Zamiaceae) by a community of the central depression of Chiapas, Mexico. Bol. Soc. Bot. Méx. 78:107-113. (1.6 Mb)

2005

• Marler, T. E., V. Lee, and C. A. Shaw. 2005. Cycad toxins and neurological diseases in Guam: Defining theoretical and experimental standards for correlating human disease with environmental toxins. HortScience 40(6):1598-1606. (1 Mb)

• Marler, T. E., V. Lee, and C. A. Shaw. 2005. Spatial variation of steryl glucosides in Cycas micronesica plants: Within- and among-plant sampling procedures. HortScience 40(6):1607-1611. (1 Mb)